PDF《28. ERITRICHIUM Schrader, Comment.》

Flora of China 16: 378C390.

1995. 28. ERITRICHIUM Schrader, Comment. Soc. Regiae Sci. Gott. 4: 186. 1820. 齿缘草属 chi yuan cao shu Plants perennial or annual, appressed strigose or villous. Leaves alternate. Cymes terminal, not branched or branched and paniculate, rarely 1-flowered. Fruiting pedicel erect or deflexed. Calyx 5-parted to base or nearly so;

lobes vertical to reflexed, enlarged in fruit. Corolla blue, light blue, or light purple, rarely yellow or white, campanulate-rotate to campanulate-tubular;

throat with appendages, rarely appendages obscure or absent;

lobes overlapping in bud, erect or divaricate at anthesis. Stamens included;

anthers orbicular, ovate to oblong. Style and stigma solitary, usually not exceeding nutlets. Gynobase flattened- fastigiate or semiglobose to

1 mm, wider than tall at base. Nutlets 4, all or some developed, turbinate to ovoid or triangular-ovoid and dorsiventrally compressed;

abaxially usually discoid, ribs or margin usually winged, dentate or with glochids. About

50 species: mainly in Asia, a few in Europe and North America;

39 species in China. 1a. Herbs annual or biennial. 2a. Corolla campanulate-rotate 24. E. heterocarpum 2b. Corolla campanulate-tubular. 3a. Stems numerous, cespitose or diffuse, branched at base or not. 4a. Stems several, cespitose;

inflorescences racemelike;

corolla light blue, light purple, or yellow, appendages obscure 28. E. tangkulaense 4b. Stems diffuse, prostrate;

flowers solitary, axillary or extra-axillary;

corolla white, appendages transverse linear 29. E. axillare 3b. Stems single or few, markedly erect, much branched above. 5a. Flowers few, widely spaced;

marginal glochids of nutlets needlelike, remote at base 27. E. acicularum 5b. Flowers numerous, crowded;

marginal glochids of nutlets lanceolate, base usually confluent into a wing. 6a. Nutlets with dense tubercles and bristles, marginal glochids ciliate, attachment scar slightly above middle adaxially 25. E. echinocaryum 6b. Nutlets glabrous or sparsely pubescent, attachment scar at middle or slightly below adaxially 26. E. thymifolium 1b. Herbs perennial. 7a. Stems slightly longer than basal leaves or not, base with crowded withered leaves and persistent petioles becoming cushionlike, or unbranched stems in leaf clusters forming rosettes. 8a. Stems single or several, basal leaves forming a rosette 38. E. villosum 8b. Stems numerous, becoming cushionlike. 9a. Corolla campanulate-tubular. 10a. Stems usually ca. as long as basal leaves;

leaf blade glabrous except for a few short hairs near apex of abaxial surface 39. E. hemisphaericum 10b. Stems longer than basal leaves;

leaf blade appressed white pubescent or spreading pilose. 11a. Stems 5C10 cm tall;

leaf blade oblanceolate, apex obtuse;

inflorescences 3C10- flowered;

calyx lobes not reflexed in fruit;

nutlets 1C1.2 mm excluding marginal glochids, marginal glochids free or nearly so at base 22. E. lasiocarpum 11b. Stems 2C5 cm tall;

leaf blade oblong to broadly elliptic or obovate, apex obtuse to acute;

inflorescences 1C4-flowered;

calyx lobes reflexed in fruit;

nutlets 1.8C2 mm excluding marginal glochids, marginal glochids confluent below middle forming a broad wing 23. E. qofengense 9b. Corolla campanulate-rotate. 12a. Marginal rib of nutlet winged or with prickles, usually without glochids. 13a. Nutlet margin winged, wing erose at margin 21. E. humillimum 13b. Nutlet margin prickly, rarely with some glochids. 14a. Leaf blade spatulate;

nutlets ca.

1 mm 19. E. pectinatociliatum 14b. Leaf blade narrowly elliptic;